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He received his bachelors's degree from Washington & Jefferson College, and his Ph.D at the University of the Witwatersrand, South Africa in 1984.
His research focuses on the hominin fossil record, during the Pliocene and early Pleistocene and the reconstruction of phylogenetic relationships through dental morphology. His most important work has been the analysis of dental microwear in order to reconstruct early hominin dietary habits.
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